Objectives

  1. Understand the main classes of antibiotics and their mechanism of action
  2. Discuss the molecular mechanism of antibiotic resistance
  3. Discuss how antibiotic resistance spreads
  4. Be a good steward of antimicrobials

Chemotherapy, caesareans and joint replacements may become too dangerous if antibiotic resistance wins.

Mechanisms of resistance

1. Restricted access to target

Antibiotic must access its target.

Permeability

Bacteria can make it so that less drug can get into cell

for example ß-lactams must cross outer membranes of gram negatices and gram negatives are intrinsically resistant to vancomysin

an example of chagning permeability is by changing porins.

Efflux pump

Bacteria can pump out drug (active efflux pump)

2. Inactivation of antibiotic

Bacteria can change the antibiotic so it does not work.

an example of this is ß-lactamase. this catalyses the ß-lactam to its inactive state so it cannot inhibit penicillin binding protein

ß-lactamase inhibitors

this is a no u to the class A ß-lactamases as it bind irreversibly to the ß-lactamases

3. Modification of target

Theis the bacteria changing the target so the mechansim of action of the antibiotic is inhibited

An example of this is a new Penicillin binding protein which does not irreversibly bind to penicillin

or vancomysin resistant peptidoglycan side chains

Target modification through mutation (fluoroquinolone)

Mutation can just develop resistance during infection

Intrinsic vs acquired resistance

Intrinsic

  • normally resistant to antimicrobial
  • chromosomally encoded
  • Examples:
    • Vancomycin and gram-negatives
    • chromosomal AmpC ß-Lactamase

Acquired

  • Bacterial normally susceptable to antimicrobials
  • Acquire resistance through gene acquisition or mutation

Spread of antimicrobial resistance

Often a route is as follows antibiotic mediated selection leads to drug resistant bacteria

drug resistance bacteria horizontally transfers resistance gene to other bacteria through 3 main ways

Gene transfer

  1. Conjugation bacterial sex

  2. transformation take up plasmid from environment

  3. transduction phages take antimicrobial resistance with them

Mobile genetic elements